Betty gets the chance to go to London to cover fashion week, bringing along Amanda and Hilda. Once there she meets up with Christina and also runs into Gio. Meanwhile, Wilhelmina joins AA – although with an ulterior motive.

When Betty goes to get her braces off, she is knocked unconcious and enters a dreamlike state where she sees how her life would’ve turned out if she’d never gotten her braces in the first place.

Betty meets a new man and wants to do a feature on him for Mode, Justin makes new friends at his acting class, and Wilhelmina is her usual ‘diva’ self despite being in hospital.

To speed along the investigation into her house fire, Betty is pressured to date an annoying fireman, Jimmy. Wilhemina tries to prove to Don, an old flame from her past, that she’s just “Wanda from the block” instead of an uber-bitch, and Amanda draws closer to Tyler, convincing Claire to hire him as an in-house model at Mode. Meanwhile, Hilda meets Bobby’s parents for the first time.

Wilhemina marginalizes Betty at Fashion Week, but Daniel has her reassigned to cover a low-priority show. There, Betty meets Marisa, one of the “10 Designers to Watch.” Meanwhile, Amanda teams up with Helen to start a new line, but both Marc and Betty refuse to show their work to Wilhelmina.

Wilhemina is livid when she lays eyes on drag queen impersonator – Wilhediva Hater. Betty and Daniel go through a rough patch as she wins an award for her blog. Daniel has the press hot on his heels. While Marc is pleased with himself when a famous director does a photo shoot with him.

Betty plans a karaoke party in her apartment building, however her building gets robbed. Marc loses something valuable. Meanwhile, Amanda tricks Daniel and Wilhelmina into attending a couples counseling session with Dr. Sparks.

Betty recalls everything that has made up her worst week in history, and also has job issues caused by taking up a new hobby. Hilda is treated to a week of luxury by Bobby, and Daniel is becoming suspicious of Mark’s actions.

Betty wants to spend some time apart from Matt, so finds him something else to keep him occupied. Meanwhile, Hilda has a tough decision to make regarding her baby, and Mark’s fling looks to be heading towards something more serious.

Ignacio prepares a combination Christmas/Hanukah dinner for the family and his new interest Jean. Betty and Hilda take the Be-Shure instant pregnancy tests. Meanwhile, Hartley hires an outsider to the fashion world named Denise Ludwig to replace Wilhemina. However Willie has her own plans for coming back. Marc helps Daniel formulate a plan to wrest control of Mead back from Hartley. Amanda helps Claire track down and meet Tyler, the son she gave up for adoption.